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What causes the "Rainbow" toning on pennies?

In a collection I just purchased there were several rolls of 1964-1966 uncirculated pennies. They were still in the original paper rolls. (Neat old papers) I opened a few for fun and they had somewhat of a "dusty" feeling to them and had darkened. I assume that was from the old paper? There were also several that had beautiful rainbow toning to them. Is this caused by the old paper rolls? Thanks
References: Multiple deals with cohodk. Also PQPeace, LordMarcoVan,
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Well, just Love coins, period.
The Penny Lady®
Collector of Early 20th Century U.S. Coinage.
ANA Member R-3147111